The World Flora Online

4. Brunellia hygrothermica Cuatrec., Fieldiana, Bot. 27: 76 (1951). Holotipo: Colombia, Cuatrecasas 17156 (F!). Ilustr.: Cuatrecasas, Fl. Neotrop. 2: 91, t. 37, 92, t. 38 (1970).

Por S. Knapp.

Árboles c. 7 m; ramitas glabras y negras cuando secas; vástagos y hojas jóvenes no vistos en el material mesoamericano, esparcidamente adpreso pelosos en el material sudamericano. Estípulas 1-1.5 mm, geminadas, triangular-oblongas, con algunos tricomas esparcidos, prontamente caducas, dejando una tenue línea interpeciolar. Hojas 30-80 cm; pecíolo 13-35 cm, glabro; eje con algunos tricomas esparcidos, adpresos, dorados, más densos en el sulco; separación entre pares de pinnas 6-8 cm; pinnas (2-)5-6 pares; estipelas bigeminadas, el par distal 1.5-2 mm, los pares basales 0.5-1 mm, todos subulados, esparcidamente adpreso pelosos; folíolos 14-23 × 6-8.5 cm, elípticos, glabros o con algunos tricomas adpresos, diminutos, a lo largo de las nervaduras en el haz, pelosos en el envés, los tricomas dorados, erectos, más densos a lo largo de las nervaduras, la base aguda a obtusa, los márgenes irregularmente crenados, algunas veces doblemente crenados, el ápice agudo; peciólulos 0-5 mm; nervadura oscura cuando seca en el envés, con c. 20 pares de nervaduras primarias, más cercanamente espaciadas en la base del folíolo. Inflorescencias 30-60 cm, difusamente ramificadas muchas veces, glabras o diminutamente adpreso pelosas, las bractéolas c. 1 mm, subuladas, prontamente caducas, los pedicelos 3-5 mm, articulados en el 1/2-1/4 inferior. Flores bisexuales, 4-5-meras; cáliz 4-5 mm de diámetro, los lobos 1.5-2 mm, triangulares, diminuta y esparcidamente dorado pelosos, los márgenes densamente pelosos, el estilo c. 1 mm, los filamentos 1.5-2 mm, densamente rojizo pelosos con tricomas crespos en la base, las anteras c. 0.5 mm, elipsoidales, basifijas, cordatas en la base. Folículos 4-5, 3-4 mm, uniforme y densamente pelosos con tricomas cortos, dorados, también con cerdas 0.5-1 mm, doradas, esparcidas, frecuentemente caducas, así que no están presentes en todos los folículos, el endocarpo cartilaginoso, rojizo; semillas c. 1.5 mm, elipsoidales, brillantes y pardo oscuro. Selvas altas perennifolias, vegetación secundaria. CR ( Herrera 4019, MO). c. 500 m. (Costa Rica, Colombia.)

Source: [
• 2
• 1
]. 

Tree 10-15 m high, the trunk 20-25 cm in diameter, with light brown bark and soft, white wood. Branchlets subterete, smooth, glabrous, brownish-green, the internodes long, only the terminal one compressed. Buds and very young opening leaves minutely ochraceous-strigose. Stipules geminate, very small, triangular-oblong, acute, 1.0-1.5 m long, soon deciduous. Leaves opposite, 1-4(-5)-jugate, very large, long-petiolate; axis 30-75 cm long, subterete, slightly striolate, narrowly sulcate above, more or less minutely, almost inconspicuously puberulous with very minute (0.05-0.10 mm long) acute, patulous or subpatulous yellowish trichomes more abundant toward the end and in the furrows, sparse toward the petiole, the interjuga 6-11 cm long, the petiole robust, glabrate, 20-40 cm long; stipels geminate, broadly corniculate, 1.0-1.5 mm long. Leaflets thinly coriaceous, large; petiolules rather robust, sulcate above, 8-15 mm long, the terminal one 28-55 cm long; blades 17-28 cm long, 8.0-13.5 cm wide, oblong-elliptic or oblong-ovate, rounded and asymmetric at the base, the terminal cuneate and symmetric, attenuate and acuminate at the apex, coarsely double-dentate-serrate, the small, callose, ascendant teeth 8-15 mm apart; green or dark green, when dry brownish-green; the midrib flat above, very conspicuous, sparsely and minutely strigulose, in part glabrous, the secondary nerves impressed, very conspicuous with an angle of divergence 45°-60°, the tertiary transversal nerves also conspicuous; whitish-green or almost white in vivo beneath, the costa thick and protuberant, the prominent secondary nerves 14-19 on each side, 8-18 mm apart, ascendent, parallel, curving near the margin, the tertiary nerves parallel, transversal, also prominent, the minor veins united into a conspicuous, more or less prominulous reticulum, the flat areoles densely whitish-papillose and with sparse, minute, acute, patulous hairs, the nervation minutely pilose with copious minute, acute, rather patulous or ascending hairs (0.05-0.10 mm long). Panicles axillary, large, long-pedunculate, 30-60 cm long and broad; peduncle robust, subterete, reddish-greenish, 12-22 cm long, glabrous or with very sparse, minute strigae; primary branching (usually subglabrate) trichotomous, the following usually dichotomous, repeatedly dividing, both spreading and ascending into a large floriferous but rather lax inflorescence, the terminal branchlets or peduncles one-flowered; branchlets thin, angular, striate, copiously and minutely strigulose or substrigulose, the hairs subappressed, acute, minute. Bracteoles above the short peduncles subulate, pilose, 0.6 mm long. Pedicels 0.5-1.0 mm long, sparsely strigulose. Flowers hermaphrodite, perfectly pentamerous, the buds angulate-globose, depressed, 1.8-2.2 mm in diameter, strigulose; calyx 2 mm long or shorter, when expanded 4.5-5.0 mm in diameter, the lobes ovate-triangular, acute, sparsely minute-strigulose outside, villous on the upper half inside, minutely tomentulose at the margin, 1.5-1.8 mm long, 1.0-1.4 mm wide; stamens ten, apparently fertile, the filament 1.5-1.8 mm long, at the lower part crisped-pilose, the anthers elliptic, cordate at the base, acute at the apex, 0.8-1.0 mm long, polliniferous; carpels five, the ovary about 0.8 mm long, ovate, hispidulo-hirsutous, biovulate, fertile, the style about 1 mm long, curved, glabrous; disc tomentose-hirsutulous. Follicles (4-) 5 maturing from a flower, round-ovoid, about 2 mm in diameter (specimens seen immature), minutely tomentulose and hispid above, the bristles 0.9 mm long; exocarp mediocre or thin; endocarp cartilagineous, oblong-ellipsoid, 1.8 mm long (immature). Seeds single, ellipsoid-globose, appressed, reddish, about 1.5 mm long.

Source: [
• 3
]. 

Restricted to the lower, coastal region of rainforest on the Pacific Coast of Colombia.

Source: [
• 3
].