The World Flora Online

Large tree, about 30 m tall. Branchlets thick, robust, quadrangular, more or less compressed, slightly striate-sulcate, densely velvety-tomentose. Young shoots, leaves and buds densely and thickly velvety, ochraceous-green or tawny. Stipules geminate, linear-triangular, thickly velvety, 3-5 mm long. Leaves opposite, large, 2-3-jugate; axis 2440 cm long, robust, inconspicuously striate, markedly sulcate above, densely velvety-tomentose, tawny, the interjuga 6-9 cm long, the petiole slightly thickened at the base, 16-23 cm long; stipels bigeminous, thickly linear-subulate, densely velvety, arcuate, the main pair 4-6 mm long, the lower smaller pair 2-3 mm long, inserted at 3-8 mm distance, deciduous, leaving a round scar. Leaflets subcoriaceous, rather rigid; petiolules 3-10 mm long, robust, tomentose, sulcate above, the terminal one 30-50 mm long; blades 12-28 cm long, 10-17 cm broad, gray-green above, green-ochraceous beneath, ovate-elliptic or obovate-elliptic, rounded, somewhat asymmetric at the base except the symmetric terminal cuneate leaflet, attenuate and subacute or obtuse at the apex, coarsely biserrate-dentate, the callous mucros 0.5-0.6 mm long, rather patulous; with broad, densely and thickly velvety midrib and tomentose secondary nerves above, for the rest pubescent with patulous, fine hairs up to 0.7 mm long, the transverse tertiary nerves and the minute venular reticulum conspicuous on the slightly rugose surface; softly tomentose beneath, the costa thick and prominent, inconspicuously striolate, rather carinate, the secondary nerves 15-20 pairs, prominent, subpatulous with angle of divergence 50°-65°, 6-18 mm apart, curvedly ascending near the margin, the tertiary nerves transversal, prominent, the minor veins united into a minute, prominulous reticulum, the main nervation densely tomentose or velvety, for the rest densely spreading, pubescent-villous. Panicles axillary, large and many-flowered, 14-26 cm long and broad, the primary branching dichasial, the lateral and terminal branches repeatedly brachiate-dichotomous, the axis obtusely angulate and striolate, densely tomentose and somewhat velvety, the branchlets densely tomentose-hirtulous, the peduncle robust, 4-6 cm long; flowers subsessile, glomerate at the end of short branchlets. Bracteoles 2-3 mm long, subulate, the inferior bracts linear, 4 mm long, densely villous. Pedicels 0.4-1.0 mm long, minutely tomentose. Female flowers mostly pentamerous, rarely tetramerous with perfect verticils; calyx about 1.5 mm long, when expanded 4-5 mm in diameter, the lobes ovate-triangular, acute, 1.7-2.0 mm long, 1.0-1.3 mm wide, densely subappressed pilose outside, sericeous inside, tomentellous at the margin and the apex; staminodia small, the filament glabrous, 0.7-0.8 mm long, the sterile anthers oblong, 0.6 mm long; carpels five, rarely four, the ovary about 1 mm long, biovulate, densely pubescent-hirsutulous, the style about 1.4 mm long, pilose toward the base; disc hirsutulous-tomentose. Follicles usually all developed, four or five maturing from a flower, crowded in a stellate arrangement, rounded, slightly compressed, 2.2-2.4 mm in diam, apiculate with the divergent stylar apiculum curved upwards, the exocarp moderately thick, densely subhirsute-tomentose, the hairs ascending, rather rigid, short and medium sized, the longest 0.3 mm long, the apicular style hairy below, about 1.5 mm long; endocarp cartilaginous, round-ellipsoid with a very obtuse insertion angle, the top rounded, 2.0 × 1.3-1.4 mm, when dry and open U-shaped. Seeds single, ellipsoid-globose, depressed, 1.6-1.7 mm in diameter, reddish-brown, lustrous.

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(Fig 40, p 96). Known only from the western slopes of the western Cordillera in Valle, Colombia, in subandean forests with heavy rainfall.

Source: [
• 1
].