The World Flora Online

Plants perennial. Stem and branches slender, hispid or villous, glabrescent. Petiole 2-4 cm; leaf blade ovate-cordate, 4-8 × 4-8 cm, membranous, 5-angular or 3-5-lobed; middle lobe larger, triangular, both surfaces scabrous, base cordate, apex acuminate. Plants monoecious. Male flowers solitary, or 3-8 in a raceme; peduncle slender, 10-15 cm; bracts leaflike, 1-2.5 cm, yellow-brown villous, 3-lobed; calyx tube tubular, elongate, ca. 2 cm; segments linear, ca. 7 mm; corolla white; segments oblong-ovate, 15-20 × 10-12 mm, ± villous; filaments ca. 0.5 mm; anthers ca. 7 mm. Female flowers solitary; pedicels 1-4 cm; ovary oblong, 10-12 × ca. 5 mm, yellow-brown villous, acute at both ends; style 5-8 mm; stigmas 3. Fruit orange, oblong-ovoid, 4-5 cm, smooth, 10-ribbed, acute at both ends. Seeds oblong, ca. 7 × 3-3.5 mm, both ends obtuse. Fl. Jul-Sep, fr. Sep-Dec.

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Climbing or creeping herb, rooting at the nodes, to 6 m long, finely scabrous and sparsely hairy or subglabrous. Probract (minute, caducous or) absent. Tendrils unbranched. Leaves: blade ovate, or triangular in outline, or 3- or 5-angled or (deeply) lobed, 4–12 cm diam., membranous, ± sparsely scabrous-pubescent on both surfaces, upper surface not bullate, cystoliths usually faint, or fine, base deeply cordate, margin finely sparsely dentate, apex acute(-acuminate), main veins 5, subpalmate; petiole 2–5 cm long. Male inflorescences: flowers solitary or severral in erect racemes 10–25 cm long, bracteate, sometimes co-axillary with a solitary flower; peduncle 4–12 cm long; bracts oblong, 15–20 mm long, (deeply) 2–5-lobed, base cuneate or rounded, sessile, without glands. Male flowers thinly grey pubescent; pedicel 3–7 cm long for solitary flowers, 5–15 mm long in the racemes, faintly articulate with receptacle; receptacle-tube narrow, but widened in upper ⅓ where containing the synandrium, 25–35 mm long, 3–4(–5) mm wide at throat, outside and inside ± pubescent, throat yellow inside; sepals linear, rarely ± lobed, 6–9 mm long, spreading to recurved, green; petals obovate, short-clawed, 20–30 by 12–15 mm, indistinctly veined, somewhat hairy, yellow at base; stamens inserted 12–15 mm below receptacle throat, filaments short, synandrium 9–10 by ca 3.5 mm, deeply included, connectives not enlarged; disc short, 3-lobed. Female flowers: perianth as in male flowers; pedicel 0.5–2(–4) cm long; ovary narrowly ellipsoid, 10–12 by 2.5–4 mm, pubescent, (faintly) 10-ribbed; style 18–20 mm long; stigmas 4–6 mm long, included; disc low, ± 3-lobed. Fruits broadly fusiform, 2.5–5 by 1.5–3 cm, beaked by receptacle remnant, orange-red, glabrescent, sharply 10-ribbed, pulp greenish (brown); fruiting pedicel 1–4 cm long. Seeds oblong, 7–8 by 2–3 by 1.5 mm, with marginal groove, faces not ornamented.

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• 4
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Forest edges, clearings, scrub, in hedges and in open fields; 0–1,000 m. altitude; flowering & fruiting throughout the year.

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• 4
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Widespread; north-eastern part of India, China, through Indochina (type), east to Sumatra, Peninsular Malaysia, Borneo, Java, Philippines, Celebes, Lesser Sunda Islands east to Flores.

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• 4
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