The World Flora Online

Herbs, perennial, to 100 cm; rhizomes absent; stolons present; corms present. Leaves emersed; petiole 5-ridged, ascending to erect, 24.5--60 cm; blade sagittate, 11.5--26.5 ´ 0.8--15 cm, basal lobes longer than remainder of blade. Inflorescences racemes, rarely panicles, of 5--17 whorls, emersed, 20--37 ´ 5--27 cm; peduncles 25--96 cm; bracts connate more than or equal to ¼ total length, lanceolate, 6.5--15 mm, delicate, not papillose; fruiting pedicels spreading, cylindric, 1.5--4.4 cm. Flowers to 3 cm diam.; sepals recurved to spreading, not enclosing flower; filaments cylindric, shorter than anthers, glabrous; pistillate flowers pedicellate, without ring of sterile stamens. Fruiting heads 0.9--1.5 cm diam; achenes oblanceoloid, abaxially keeled, 1.2--2.5 ´ 0.8--1.6 mm, beaked; faces tuberculate, wings absent, glands 0--1; beak lateral, erect, 0.1--0.6 mm.

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5. Sagittaria longiloba Engelm. ex J.G. Smith, N. Amer. Sagittaria 16, pl. 11 (1894). Lectotipo (designado por Bogin, 1955): Estados Unidos, Gunnison Expedition 49 (NY).

Por A. Lot H. y A. Novelo R.

Rizoma corto y grueso, comúnmente con estolones. Hojas erectas; pecíolos hasta 55 cm, subcilíndricos, acanalados; láminas 13-26 x 2-2.5, lanceolado triangulares, la base sagitada con lóbulos basales casi el doble del tamaño de la lámina, el ápice agudo. Inflorescencia simple; escapos hasta 70 cm, erectos, triangulares desde la base y entre los verticilos, acanalados; verticilos 4-5, cada uno con (2-)3(-4) flores. Brácteas 0.3-0.5 mm, connatas en la base. Flores estaminadas con pedicelos 1.5-2.8 cm, erectos; sépalos c. 6 mm; pétalos casi el doble del largo de los sépalos, blancos; estambres c. 20; filamentos lineares. Flores pistiladas en varios de los verticilos inferiores; pedicelos 1.5-4 cm, erectos, no engrosados en fruto; sépalos c. 5 mm, reflexos; pétalos casi el doble del largo de los sépalos, blancos. Cabezuelas fructíferas c. 1 cm, esféricas; aquenios 2-2.3 x 0.9-1.2 mm, obovoides, con 1 costilla, dorsal y ventralmente alados, con 1 glándula alargada c. 1 mm; pico estilar 0.2-0.3 mm. Pantanos, zanjas temporales. N (Clewell 3207, MO). 100-300 m. (S. Estados Unidos, México, Nicaragua, Venezuela.)

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Sagittaria longiloba Engelm. ex J.G. Sm., N. Amer. Sagittaria 16. 1894; S. greggii J.G. Sm.

Hierbas perennes, glabras, hasta 1 m de alto; estolones hasta 15 cm de largo y 0.6 cm de diámetro, cormos ca 3 cm de diámetro. Hojas emergentes, sagitadas, 11.5–26.5 cm de largo y 0.8–15 cm de ancho, ápice agudo a obtuso, enteras, con 5–15 nervios, lobos basales 6–19 cm de largo, enteros, separados entre sí 2.2–28 cm, con ca 6 nervios; pecíolos aplanados, 24.5–60 cm de largo, con una vaina 5.2–13 cm de largo en la base. Inflorescencia un escapo con 5–17 verticilos, 20–37 cm de largo y 5–27 cm de ancho, los verticilos con 3 flores, los más inferiores raramente ramificados, los inferiores pistilados, los superiores estaminados, pedúnculos teretes, 25–96 cm de largo, acostillados, brácteas subuladas, 0.8–4.5 cm de largo, delicadas a ligeramente gruesas, glabras, con el ápice largamente atenuado, pedicelos patentes a ascendentes cuando en flor y fruto, cilíndricos, 0.5–4.5 cm de largo y 0.3–0.9 mm de diámetro, flores unisexuales; sépalos libres, reflexos a patentes, 5–9 mm de largo y 2.4–6 mm de ancho, verdes; pétalos unguiculados, libres, patentes, 5–15 mm de largo y 3.7–14 mm de ancho, blancos, la uña 1.4–3 mm de largo; estambres 12–16, libres, anteras 1.5–2.2 mm de largo y 0.3–1 mm de ancho, filamentos lineares, 1–3 mm de largo, glabros; carpelos numerosos y libres. Frutos aplanados, rostrados, 1.2–2.5 mm de largo y 0.8–1.6 mm de ancho, caras tuberculadas, con un ala y a veces con 1 glándula, ésta linear a semicircular, dispuesta a lo largo del ala, rostro lateral, erecto, 0.1–0.6 mm de largo.

Localmente abundante en aguas poco profundas y en sitios inundados, en Granada y Matagalpa; 30–500 m; fl y fr feb–jun; Grijalva 129, Haynes 8300; Estados Unidos (Nebraska y California) hasta Nicaragua.

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Perennial with rhizomes and/or runners with corms. Leaves emersed, sagittate, with linear to ovate-triangular blades 3-15 x 1.3-10 cm, the linear to lanceolate basal lobes 8-21 X 0.6-7 cm, always longer than and commonly twice as long as the blades, the petioles 35-75 cm long. Scapes 35-140 cm long, with 4-12 whorls of flowers, commonly branching at the lowest whorl. Bracts linear, elongate, 0.8-5 cm long, joined at the base, the free ends often reflexed. Pedicels 1-3.5 cm long, ascending. Sepals 0.4-0.7 cm long, reflexed. Petals white, ca. twice the length of the sepals. Stamens 15- [infinity], the glabrous, linear filaments 2-3 mm long, exceeding the nearly linear anthers. Mature receptacle 0.7-1.2 cm in diameter; achenes obovate, 1.2-2.3 x 0.6-1.3 mm, the narrow dorsal wing to 0.3 mm wide, the ventral wing nearly obsolete, the faces commonly 1-winged, the short, triangular, laterally inserted beak to 0.15 mm long, or obsolete.

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Perennial herbs, from stolons or corms, glabrous, to 100 cm tall; stolons to 15 cm long, to 0.6 cm wide; corms, cm long, ca. 3 cm diam. Leaves emersed, petiolate, the petioles flattened, 24.5-60 cm long, 2-8.5 mm wide, the basal sheath 5.2-13 cm, the blades pale green, sagittate, 11.5-26.5 cm long, 0.8-15 cm wide, with 5-15 veins, the apex acute to obtuse; basal lobes 6-19 cm long, the tips separated by 2.2-28 cm, with ca. 6 veins. Inflorescence a simple or rarely paniculate scape of 5-17 whorls, emersed, 20-37 cm long, 5-27 cm wide, the whorls with 3 flowers; peduncles terete, 25-96 cm long, 2-7.5 mm diam.; staminate bracts united, lanceolate, delicate, 6.5-10 mm long, the apex acute; carpellate bracts united, lanceolate, delicate, 1-1.5 cm long, the apex acute; staminate pedicels spreading, cylindric, 2-2.5 cm long, 0.2-0.5 mm diam.; carpellate pedicels spreading in flower and fruit, cylindric, 1.5-1.4 cm long, 0.4-0.8 mm diam. Staminate flowers without sterile carpels, the sepals reflexed to spreading, 5-9 mm long, 3-6 mm wide, the petal clawed, 5-12 mm long, 3.7-14 mm wide, the claws 1.4-3 mm long, the stamens 12-16, the filaments glabrous, cylindric, 1-3 mm long, ca. 0.1 mm wide, the anthers linear, 1.5-2.2 mm long, 0.3-1 mm wide, the apex obtuse; carpellate flowers without ring of sterile stamens, the sepals reflexed to spreading in flower and fruit, 5.5-7.5 mm long, 24-3.5 mm wide, the petals clawed, 6-15 mm long, 4-10 mm wide, the claw ca. 2 mm long. Fruit aggregate 0.9-1.5 cm diam.; achenes oblanceolate, beaked, with keel, 1.2-2.5 mm long, 0.8-1.6 mm wide, the faces tuberculate, with 0-1 glands, the glands linear to cresent, the beak lateral, erect, 0.1-0.6 mm long.

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Nebraska and California, south to Mexico, but absent in the Rocky Mountains and the Great Basin. A common plant of the southern Great Plains and Mexico, in the shallow waters of sloughs, swamps, ponds, and ditches. Collections examined from Arizona, California, Colorado, Kansas, Nebraska, Oklahoma, and Texas in the United States, and Guanjuato, Michoacan, Oaxaca, Queretaro, Sinaloa, Sonora, and Tamaulipas in Mexico.

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Western United States of America, Mexico, and Nicaragua.

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