The World Flora Online

Arbres, arbustes, ou lianes avec huiles essentielles. Feuilles alternes, pétiolées, sans stipules, simples ou composées-digitées. Inflorescences axillaires ou terminales, corymbiformes ou paniculées, pourvues ou non des bractées. Fleurs unisexuées ou bisexuées, actinomorphes, périgone à tépales libres ou légèrement soudés à la base, en deux verticilles 3-5-mères à préfloraison valvaire ou en 1 verticille 4-8-mère à préfloraison imbriquée; étamines 3-5, en 1 verticille, opposées aux tépales externes si le périgone est double; anthères à 2 thèques s'ouvrant par des valves (1 par thèque) introrses ou latérales; staminodes en 1 ou 2 verticilles et glanduliformes, ou absents: ceux du verticille externe par paire à la base des étamines, ceux du verticille interne alternant avec les étamines; ovaire infère, uniloculaire avec un style unique, à 1 ovule pendant et anatrope. Fruits secs, non ailés ou à 2-4 ailes longitudinales ou à 2 ailes apicales. Graines solitaires, exalbuminées; embryon droit; cotylédons plan-convexes ou spiralés.

Herbes , arbustes ou lianes.'Feuilles'exstipulées, alternes, simples ou composées-digitées.'Inflorescences'axillaires ou terminales, pédonculées, corymbiformes ou paniculées, pourvues ou non de bractées.'Fleurs'toutes ☿ ou les unes ☿ et les autres ♂♀ par avortement, régulières ; périgone à tépales libres ou légèrement soudés à la base, en 2 verticilles 3-5-mères à préfloraison valvaire ou en 1 verticille 4-8-mère à préfloraison imbriquée; étamines 3-5 en 1 verticille, opposées aux tépales externes si le périgone est double; anthères à 2 thèques s'ouvrant par valves introrses ou latérales; staminodes glanduliformes en 1 ou 2 verticilles, ou absents : ceux du verticille externe par paire à la base des étamines, ceux du verticille interne alternant avec les étamines; ovaire infère, 1-loculaire; ovule 1, pendant et anatrope.'Fruits'secs ou drupacés, à 2-4 ailes longitudinales ou à 2 ailes terminales-apicales, ou non ailés et enfermés dans une cupule renflée.'Graines solitaires, exalbuminées; embryon droit; cotylédons plans-convexes ou spiralés.\n\t\t\tQuatre genres et ± 60 espèces dans les régions tropicales.

Stamens 3–5 in a single whorl, when the perianth is double opposite the outer segments, often 4; anthers 2-celled, opening introrsely or laterally by 2 valves; staminodes gland-like, in one or two whorls outside the stamens, or absent, those of the outer whorl in pairs at the base of the filaments, those of the inner whorl alternate with the stamens Ovary single, inferior, 1-celled; ovule solitary, pendulous Calyx with 3–5 valvate subequal segments in two whorls or rarely 4–8 in one whorl Fruit dry, more or less ribbed, either with 2–4 wings on the body or with two terminal wings formed by enlarged perianth-segments, or wingless but enclosed in the inflated receptacle Seed solitary, without endosperm; testa leathery; embryo straight; cotyledons large, plano-convex or flat and twisted around the radicle Trees or shrubs, sometimes scandent, with alternate, simple or digitately compound leaves; stipules absent Flowers hermaphrodite, or monoecious or polygamous by abortion, actinomorphic, arranged in axillary corymbose or paniculate cymes, bracteate or not Ovary single, inferior, 1-celled; ovule solitary, pendulous Calyx with 3–5 valvate subequal segments in two whorls or rarely 4–8 in one whorl Fruit dry, more or less ribbed, either with 2–4 wings on the body or with two terminal wings formed by enlarged perianth-segments, or wingless but enclosed in the inflated receptacle Seed solitary, without endosperm; testa leathery; embryo straight; cotyledons large, plano-convex or flat and twisted around the radicle Trees or shrubs, sometimes scandent, with alternate, simple or digitately compound leaves; stipules absent Flowers hermaphrodite, or monoecious or polygamous by abortion, actinomorphic, arranged in axillary corymbose or paniculate cymes, bracteate or not

Seed solitary, without endosperm; embryo straight; cotyledons plano-convex, ± lobed, or flattened and spirally twisted Leaves alternate, exstipulate, simple, digitately compound or 3–5-lobed Inflorescences of axillary, rarely terminal, corymbose or paniculate cymes, pedunculate, bracteate or ebracteate Flowers regular, bisexual, or unisexual by abortion and the plants monoecious or polygamous, rarely dioecious Perianth segments subequal, free or shortly united below, in two 3–5-merous, valvate whorls or in one 4–8-merous, imbricate whorl Stamens 3–5 in one whorl and opposite the outer segments when perianth is double, or (2)4(7) when perianth is single; anthers 2-thecous, dehiscing introrsely or laterally by 2 flap-like valves Staminodes absent or ± gland-like, 1–2-whorled, usually outside the stamens; outer whorl paired at the base of the stamens, inner whorl alternating with the stamens Ovary inferior, 1-locular; ovule solitary, pendulous, anatropous Fruit dry indehiscent, often winged, longitudinally ribbed or with lateral wings (Illigera), or with 2 apical wings formed by accrescent perianth segments (Gyrocarpus), or wingless and enclosed in an inflated capsule (Hernandia) Trees, shrubs or lianes, with aromatic oils in stems and leaves Leaves alternate, exstipulate, simple, digitately compound or 3–5-lobed Inflorescences of axillary, rarely terminal, corymbose or paniculate cymes, pedunculate, bracteate or ebracteate Flowers regular, bisexual, or unisexual by abortion and the plants monoecious or polygamous, rarely dioecious Perianth segments subequal, free or shortly united below, in two 3–5-merous, valvate whorls or in one 4–8-merous, imbricate whorl Stamens 3–5 in one whorl and opposite the outer segments when perianth is double, or (2)4(7) when perianth is single; anthers 2-thecous, dehiscing introrsely or laterally by 2 flap-like valves Staminodes absent or ± gland-like, 1–2-whorled, usually outside the stamens; outer whorl paired at the base of the stamens, inner whorl alternating with the stamens Ovary inferior, 1-locular; ovule solitary, pendulous, anatropous Fruit dry indehiscent, often winged, longitudinally ribbed or with lateral wings (Illigera), or with 2 apical wings formed by accrescent perianth segments (Gyrocarpus), or wingless and enclosed in an inflated capsule (Hernandia) Trees, shrubs or lianes, with aromatic oils in stems and leaves

Trees, shrubs, or scandent lianas. Leaves simple or palmately compound, petiolate, circinate and scandent in part, estipulate. Flowers bisexual, unisexual, or polygamous, actinomorphic, in axillary or terminal corymbs or cymose panicles, with bracts or not. Outer tepals (sepals) 3-5. Inner tepals (petals) similar to outer ones. Stamens 3-5; filament bases with appendages on external sides or not; anthers 2-celled; cells valvate. Ovary inferior, 1-loculed, 1-ovuled; ovule pendulous. Drupe ± costate, broadly 2-4-winged or enclosed in an inflated cupule and wings absent. Seed 1, exalbuminous, coat leathery.

Seed solitary, without endosperm, enclosed in woody endocarp; embryo straight; cotyledons large, plano-convex, ± lobed, or flattened and spirally twisted Monoecious trees or shrubs or lianes with bisexual flowers, with aromatic oils in stems and leaves Flowers usually small, greenish yellow, cream-coloured or white in axillary, peduncled, corymbose or panicled, bracteate or ebracteate cymes, bisexual or polygamous by abortion, regular Leaves alternate, exstipulate, simple or palmately compound, sometimes with small pellucid dots, 3–7-nerved at or a little above the base Stamens 3–5 in a single whorl, opposite the outer perianth- segments when the perianth is double, or 2–7 but generally 4 when the perianth is single; anthers 2-thecous, dehiscing introrsely or laterally by 2 flap-like valves Perianth superior; segments free or shortly united below, in two valvate 3–5-merous whorls, or more rarely in one imbricate 4–8-merous whorl Ovary inferior, 1-locular; ovule solitary, pendulous, anatropous Staminodes ± glandular, in 1 or 2 whorls usually outside the stamens, or absent; those of the outer whorl in pairs at the base of the stamens, those of the inner whorl alternate with the stamens Fruit dry, often longitudinally ribbed, sometimes with 2–4 longitudinal wings (Illigera), or with 2 terminal wings formed by accrescent perianth-segments (>i>Gyrocarpus), or wingless and enclosed in an inflated capsule (>i>Hernandia) Monoecious trees or shrubs or lianes with bisexual flowers, with aromatic oils in stems and leaves Flowers usually small, greenish yellow, cream-coloured or white in axillary, peduncled, corymbose or panicled, bracteate or ebracteate cymes, bisexual or polygamous by abortion, regular Leaves alternate, exstipulate, simple or palmately compound, sometimes with small pellucid dots, 3–7-nerved at or a little above the base Stamens 3–5 in a single whorl, opposite the outer perianth- segments when the perianth is double, or 2–7 but generally 4 when the perianth is single; anthers 2-thecous, dehiscing introrsely or laterally by 2 flap-like valves Perianth superior; segments free or shortly united below, in two valvate 3–5-merous whorls, or more rarely in one imbricate 4–8-merous whorl Ovary inferior, 1-locular; ovule solitary, pendulous, anatropous Staminodes ± glandular, in 1 or 2 whorls usually outside the stamens, or absent; those of the outer whorl in pairs at the base of the stamens, those of the inner whorl alternate with the stamens Fruit dry, often longitudinally ribbed, sometimes with 2–4 longitudinal wings (Illigera), or with 2 terminal wings formed by accrescent perianth-segments (>i>Gyrocarpus), or wingless and enclosed in an inflated capsule (>i>Hernandia)