Plants in medium-sized to large wefts, often extensive sheets. Stems creeping to ascending or erect, sympodial or monopodial, irregularly branched to regularly pinnate, innovations often ascending-arching; paraphyllia sometimes present, branched, cells elongate; pseudoparaphyllia foliose; rhizoids few, at base of stems and apices of attenuate branches. Stem leaves very broadly ovate to lanceolate; margins serrate to serrulate, sometimes nearly entire; apex acuminate or sometimes acute to rounded; costa usually double, 1/4-2/3 leaf length, sometimes single or nearly ecostate; alar cells sometimes differentiated; medial laminal cells narrowly elliptic to linear-flexuose, prosenchymatous. Branch leaves usually smaller, narrower; margins more sharply toothed; costa stronger. Sexual condition dioicous. Seta elongate, smooth. Capsule usually inclined to horizontal; peristome double; exostome usually reticulate proximally, papillose distally; endostome basal membrane usually high, segments broad, perforate along keel by narrow slits or broadly elliptic gaps, cilia 1-4 (absent in Leptohymenium). Calyptra cucullate, smooth, naked. Spores spheric, finely papillose to nearly smooth. SELECTED REFERENCES Andrews, A. L. 1954. Taxonomic notes. XII. The families Rhytidiaceae and Hylocomiaceae. Bryologist 57: 251-261. Noguchi, A. 1972. On the delimitation of the genera of Hylocomiaceae and Rhytidiaceae. J. Hattori Bot. Lab. 35: 155-168. Hedenas, L. 2004. Morphological and anatomical evidence suggest that 'Hylocomiaceae' taxa belong to at least two clades. J. Bryol. 26: 125-135. Rohrer, J. R. 1985. A generic revision of the Hylocomiaceae. J. Hattori Bot. Lab. 59: 241-278. Rohrer, J. R. 1985b. A phenetic and phylogenetic analysis of the Hylocomiaceae and Rhytidiaceae. J. Hattori Bot. Lab. 59: 185-240. SELECTED REFERENCES Andrews, A. L. 1954. Taxonomic notes. XII. The families Rhytidiaceae and Hylocomiaceae. Bryologist 57: 251-261. Noguchi, A. 1972. On the delimitation of the genera of Hylocomiaceae and Rhytidiaceae. J. Hattori Bot. Lab. 35: 155-168. Hedenas, L. 2004. Morphological and anatomical evidence suggest that 'Hylocomiaceae' taxa belong to at least two clades. J. Bryol. 26: 125-135. Rohrer, J. R. 1985. A generic revision of the Hylocomiaceae. J. Hattori Bot. Lab. 59: 241-278. Rohrer, J. R. 1985b. A phenetic and phylogenetic analysis of the Hylocomiaceae and Rhytidiaceae. J. Hattori Bot. Lab. 59: 185-240. Plants in medium-sized to large wefts, often extensive sheets. Stems creeping to ascending or erect, sympodial or monopodial, irregularly branched to regularly pinnate, innovations often ascending-arching; paraphyllia sometimes present, branched, cells elongate; pseudoparaphyllia foliose; rhizoids few, at base of stems and apices of attenuate branches. Stem leaves very broadly ovate to lanceolate; margins serrate to serrulate, sometimes nearly entire; apex acuminate or sometimes acute to rounded; costa usually double, 1/4-2/3 leaf length, sometimes single or nearly ecostate; alar cells sometimes differentiated; medial laminal cells narrowly elliptic to linear-flexuose, prosenchymatous. Branch leaves usually smaller, narrower; margins more sharply toothed; costa stronger. Sexual condition dioicous. Seta elongate, smooth. Capsule usually inclined to horizontal; peristome double; exostome usually reticulate proximally, papillose distally; endostome basal membrane usually high, segments broad, perforate along keel by narrow slits or broadly elliptic gaps, cilia 1-4 (absent in Leptohymenium). Calyptra cucullate, smooth, naked. Spores spheric, finely papillose to nearly smooth.General Information
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Flora of North America @ efloras.org
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