Shrubs or trees (deciduous); (thorny) spines absent: glabrous or glabrate. Stems erect; profusely branched. Leaves alternate, spirally arranged, (minute, scalelike); venation pinnate; stipules present (caducous, scalelike); petiole absent. Inflorescences axillary, racemose, corymbose, or flowers solitary; pedunculate; bud scales usually persistent; bracts usually absent, sometimes present. Pedicels present. Flowers bisexual, actinomorphic, rotate to crateriform, campanulate, or urceolate; perianth and androecium hypogynous; sepals persistent, 4, distinct or connate basally; petals 4, attached directly to receptacle, imbricate, distinct, equal; intrastaminal nectary-discs or glands usually present; stamens 8; filaments distinct, glabrous or pubescent; anthers dehiscing by longitudinal slits; pistil 1; ovary 1-carpellate, 2-locular; placentation axile; ovules anatropous, bitegmic; style 1 (straight, short and thick); stigma 1, capitate, unlobed. Fruits: berries, indehiscent (spheroidal). Seeds (1 or) 2-4, blackish; not arillate; endosperm scanty or absent, sometimes a persistent perisperm present; cotyledons incumbent. x = 11. Highly branched thorny shrubs or small trees to ca. 10 m tall; branches green, glabrous to puberulent, ending in thorns; leaves alternate, scale-like to lanceolate, (1-) 1.5-2.2(-3) mm long, caducous; inflorescence lateral, umbelliform to racemose, 5-20 (or more)-flowered; flowers perfect, actino morphic; sepals 4, free, green; petals 4, free, white to creamy yellow to greenish; stamens 8, the filaments quadrangular in cross section, thickened at or slightly below the middle; anthers versatile, the dehiscence longitudinal; ovary superior, stipitate, 2-celled; ovules numerous, axile; fruit a globose berry with a thin rostellum; seeds 2-4, cochleate. SELECTED REFERENCE Holmes, W. C., K. L. Yip, and A. E. Rushing. 2008. Taxonomy of Koeberlinia (Koeberliniaceae). Brittonia 60: 171-184. SELECTED REFERENCE Holmes, W. C., K. L. Yip, and A. E. Rushing. 2008. Taxonomy of Koeberlinia (Koeberliniaceae). Brittonia 60: 171-184. Shrubs or trees (deciduous); (thorny) spines absent: glabrous or glabrate. Stems erect; profusely branched. Leaves alternate, spirally arranged, (minute, scalelike); venation pinnate; stipules present (caducous, scalelike); petiole absent. Inflorescences axillary, racemose, corymbose, or flowers solitary; pedunculate; bud scales usually persistent; bracts usually absent, sometimes present. Pedicels present. Flowers bisexual, actinomorphic, rotate to crateriform, campanulate, or urceolate; perianth and androecium hypogynous; sepals persistent, 4, distinct or connate basally; petals 4, attached directly to receptacle, imbricate, distinct, equal; intrastaminal nectary-discs or glands usually present; stamens 8; filaments distinct, glabrous or pubescent; anthers dehiscing by longitudinal slits; pistil 1; ovary 1-carpellate, 2-locular; placentation axile; ovules anatropous, bitegmic; style 1 (straight, short and thick); stigma 1, capitate, unlobed. Fruits: berries, indehiscent (spheroidal). Seeds (1 or) 2-4, blackish; not arillate; endosperm scanty or absent, sometimes a persistent perisperm present; cotyledons incumbent. x = 11. Highly branched thorny shrubs or small trees to ca. 10 m tall; branches green, glabrous to puberulent, ending in thorns; leaves alternate, scale-like to lanceolate, (1-) 1.5-2.2(-3) mm long, caducous; inflorescence lateral, umbelliform to racemose, 5-20 (or more)-flowered; flowers perfect, actino morphic; sepals 4, free, green; petals 4, free, white to creamy yellow to greenish; stamens 8, the filaments quadrangular in cross section, thickened at or slightly below the middle; anthers versatile, the dehiscence longitudinal; ovary superior, stipitate, 2-celled; ovules numerous, axile; fruit a globose berry with a thin rostellum; seeds 2-4, cochleate.General Information
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Literature
Source: [
Flora of North America @ efloras.org
LiteratureBrittonia Journal
General Information